The meaning of "The smile comprehended" in the paragraph would be made more explicit if the clause were restated as...

Little Montenegro! He lifted up the words and nodded at them--with his smile. The smile comprehended Montenegro's troubled history and sympathized with the brave struggles of the Montenegrin people. It appreciated fully the chain of national circumstances which had elicited this tribute from Montenegro's warm little heart. My incredulity was submerged in fascination now; it was like skimming hastily through a dozen magazines.

The meaning of "The smile comprehended" in this paragraph would be made more explicit if the clause were restated as?

A) The smile is understood

B) Gatsby's smile comprehended

C) The smile showed that Gatsby comprehended

D) The smile helped Nick to comprehend

E) The smile fully grasped