In this chapter of White Fang, what effect does the conflict with Lip-lip have on White Fang?

Chapter Two?

But the effect upon White Fang was not to cow him. Though he suffered most of the damage and was always defeated, his spirit remained unsubdued. Yet a bad effect was produced. He became malignant and morose. His temper had been savage by birth, but it became more savage under this unending persecution. The genial, playful, puppyish side of him found little expression. He never played and gambolled about with the other puppies of the camp. Lip-lip would not permit it. The moment White Fang appeared near them, Lip-lip was upon him, bullying and hectoring him, or fighting with him until he had driven him away.

The effect of all this was to rob White Fang of much of his puppyhood and to make him in his comportment older than his age. Denied the outlet, through play, of his energies, he recoiled upon himself and developed his mental processes. He became cunning; he had idle time in which to devote himself to thoughts of trickery. Prevented from obtaining his share of meat and fish when a general feed was given to the camp-dogs, he became a clever thief. He had to forage for himself, and he foraged well, though he was oft-times a plague to the squaws in consequence. He learned to sneak about camp, to be crafty, to know what was going on everywhere, to see and to hear everything and to reason accordingly, and successfully to devise ways and means of avoiding his implacable persecutor.